I discussed the holes with Pete Lovell and even he didn’t have a preference, saying there seems good and bad reasons for both options !
Here’s my conclusion…
The flow via the big ends will be (within reason) the same with or without the holes as it is dictated largely by the pump.
Without holes, the oil will be forced to exit out of the sides of the big ends and be flung all around the crank cases.
This oil fling will do a good job of cooling the engine and lubricating everything in there.
If the flow remains close to the same, but with the holes blocked, it stands to reason that the pressure will be higher.
I therefore run without the holes in both my engines.